:root {
  --ink: #2a2117;
  --muted: #625342;
  --paper: #f4e8cc;
  --paper-2: #fffaf0;
  --brass: #8c6726;
  --accent: #b85e32;
  --line: #bfa06a;
}

* { box-sizing: border-box; }

body {
  margin: 0;
  color: var(--ink);
  background: radial-gradient(circle at 15% 10%, #fff8e9, transparent 42%), var(--paper);
  font-family: Georgia, "Times New Roman", serif;
  line-height: 1.65;
}

.paper {
  width: min(980px, 92vw);
  margin: 32px auto;
  padding: clamp(24px, 6vw, 68px);
  background: rgba(255, 250, 235, .72);
  border: 1px solid var(--line);
  box-shadow: 0 18px 50px rgba(65, 45, 19, .16);
}

.top-nav { color: var(--muted); font-size: .92rem; text-align: right; }
.top-nav a, footer a, a { color: var(--brass); }

header { max-width: 760px; margin: 28px auto 42px; text-align: center; }
.eyebrow { color: var(--brass); font: 700 12px/1.2 ui-monospace, SFMono-Regular, Menlo, monospace; letter-spacing: .18em; text-transform: uppercase; }
h1 { margin: 12px 0 8px; font-size: clamp(2.4rem, 7vw, 4.6rem); line-height: 1; }
h2 { margin: 38px 0 10px; color: var(--brass); font-size: 1.55rem; border-bottom: 1px solid var(--line); padding-bottom: 6px; }
.lead, .scientific { color: var(--muted); font-style: italic; font-size: 1.15rem; }
.scientific { margin: 0; font-size: 1.05rem; }
.badge { display: inline-block; padding: 4px 10px; border: 1px solid var(--brass); border-radius: 999px; color: var(--brass); font: 700 11px/1.2 ui-monospace, SFMono-Regular, Menlo, monospace; letter-spacing: .08em; text-transform: uppercase; }
.privacy, .conservation, .cta-panel { margin: 24px 0; padding: 18px 20px; background: rgba(184, 94, 50, .08); border-left: 4px solid var(--accent); }
.conservation { background: rgba(140, 103, 38, .08); border-left-color: var(--brass); }
.cta-panel { text-align: center; }
.cta { display: inline-block; margin-top: 8px; padding: 12px 20px; color: #fff8ed; background: var(--accent); border-radius: 5px; text-decoration: none; font-weight: 700; }
.muted { color: var(--muted); }
ul { padding-left: 24px; }
.species-grid { display: grid; grid-template-columns: repeat(auto-fit, minmax(220px, 1fr)); gap: 12px; }
.species-card { display: flex; flex-direction: column; gap: 2px; padding: 16px; background: rgba(230, 210, 167, .42); border: 1px solid rgba(140, 103, 38, .45); color: var(--ink); text-decoration: none; }
.species-card:hover, .species-card:focus-visible { outline: 3px solid rgba(184, 94, 50, .42); outline-offset: 2px; }
.species-card strong { color: var(--brass); font-size: 1.08rem; }
.species-card em, .species-card span { color: var(--muted); font-size: .9rem; }
footer { margin-top: 42px; color: #6d5a43; font-size: .9rem; text-align: center; }

@media (max-width: 640px) {
  .paper { width: min(94vw, 980px); margin: 16px auto; }
  .top-nav { text-align: center; }
}